Neritopsines are grazers on algal spores, diatoms, and detritus. Some freshwater species such as Theodoxus are also carnivorous, feeding on aquatic insect larvae. Terrestrial representatives feed on detritus, algal spores, moss, and lichens. Deep sea and vent taxa are likely detritivores as well.
Foraging behavior for marine intertidal forms is closely tied to both light and tidal cycles. Many intertidal species move only when awash, and not when full exposed to aerial conditions or when completely submerged. Movement at low tide at night is often common. Many intertidal species also rest at a higher level in the intertidal zone and feed at a lower one.
Predators of neritopsines include crabs, fishes, and other predatory gastropods. Several tropical crab species have large, heavy claws that can effortlessly crush the shells of intertidal nerite species. Tropical reef fishes are also well equipped to both remove and crush nerites. Predatory gastropods are also common predators on neritopsine species, and several species have escape responses to the approach of a carnivorious species.
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