Semicircular Canals

The three semicircular canals project in three different planes at nearly right angles to each other. Each canal contains an inner extension of the membranous labyrinth called a semicircular duct, and at the base of each duct is an enlarged swelling called the ampulla. The crista ampullaris, an elevated area of the ampulla, is where the sensory hair cells are located. The processes of these cells are embedded in a gelatinous membrane, the cupula (fig. 10.15), which has a higher density than that of the surrounding endolymph. Like a sail in the wind, the cupula can be pushed in one direction or the other by movements of the endolymph.

The endolymph of the semicircular canals serves a function analogous to that of the otolithic membrane—it provides inertia so that the sensory processes will be bent in a direction opposite to that of the angular acceleration. As the head rotates to the right, for example, the endolymph causes the cupula to be bent toward the left, thereby stimulating the hair cells. Hair cells in the anterior semicircular canal are stimulated when doing a somersault, those in the posterior semicircular canal are stimulated when performing a cartwheel, and those in the lateral semicircular canal are stimulated when spinning around the long axis of the body.

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